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Sipuncula
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This phylum's 'place'
relative to other phyla:
Selected taxa within this taxon:
none (4 classes)

350 species

Sipunculids are burrowing worms that share many similarities with annelids: possess a trochophore larvae and similar musculature, excretory, and nervous systems (and similar ecologically with echiurans).  Also closely related based on molecular data.

But sipunculids lack segementation and setae
 

Defining characteristics
 


 
 
 

Why are sipunculids not considered annelids?  More broadly, is the grouping of animals into phyla purely arbitrary (is the concept of 'body plan' subjective).  Should classification schemes be designed for convience as well as reflect evolutionary relationships?

While evolution does not produce an unbroken continuum of organisms, some "breaks" (branches) are wider (further apart) than others.

In addition, some zoologists have argued that the sipunculans more closely related to the molluscs than to annelids, based on similarity early in development (arrangement of cells in the embryo, known as the "molluscan cross" and in sipunculid larvae a muscular creeping foot)

Fossil  of Sipuncula are typiclly rare with one possible exception.  Hyoliths (Paleozoic) are conical shells with hinged lids.  In a few species the intestine has been fossilized and is looped and coiled as in living sipunculans. A few living sipunculans secrete a calcified cuticular plate, the anal shield.

 
 
 
 
 
 
 
Lecture Sources:
  • Pechenik. Jan A.  2000. Biology of the Invertebrates.  McGraw-Hill, New York.


 
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